Folic acid (FA) food fortification and increased periconceptional FA supplementation have increased maternal folate status (1). High FA intake and maternal folate are associated with increased risk of developing gestational diabetes mellitus (GDM) (2). We hypothesise that the mechanism of FA action involves placental hormone dysregulation of maternal insulin resistance.
Maternal folate, human placental lactogen (hPL) and placental growth hormone (GH2) were measured in early-gestation blood samples taken pre- (2005-2008, n=1164) and post- (2015-2018, n=1300) FA fortification. Furthermore, early to mid-gestation (n=66, 6-16 weeks’) human placental explants were treated in vitro with increasing doses of FA (0 nM (deficiency), 40 nM (adequate), 2000 nM (physiological excess)) to establish direct effects on placental hormone (hPL and GH2) secretion and folate receptor 1 (FOLR1) expression.
GDM incidence increased from 5% pre- to 15.2% post- FA fortification. Compared to pre-fortification, women post-fortification had higher serum folate (18%, p<0.001), red cell folate (RCF; 259%, p<0.001), hPL (29%, p<0.0001) and GH2 (13%, p=0.01). RCF increased GDM risk (RR: 1.34 95% CI: 1.01-1.78, p=0.04). In vitro, compared to adequate treatment, FA deficiency resulted in higher hPL (29%, p<0.005), whilst FA excess increased placental secretion of hPL (24%, p<0.05) and GH2 (26%, p<0.05). A similar U-shaped response was seen with FOLR1, where increased expression was measured in response to FA deficiency (40%, p=0.04) and excess (54%, p=0.0004) compared to adequate FA.
Excess FA dysregulates placental hormones that regulate maternal insulin resistance in vivo and in vitro, providing a novel mechanism linking FA and GDM. Interestingly, we provide evidence that the effects of excess FA parallel those of deficiency, suggesting FA metabolism is dysregulated in high-FA conditions, increasing concerns about excess intake in pregnancy. Given nearly 20% of Australian pregnancies are now affected by GDM, it is imperative to understand its pathogenesis.